David W. Greenfield
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E-mail: greenfie@hawaii.edu

My research focuses on the systematics and ecology of coral-reef fishes. Systematic studies deal mainly with three families: Batrachoididae (toadfishes), Gobiidae (gobies), and Holocentridae (soldierfishes). Ecological studies are centered on assemblage structure in various habitats and between oceans.

A SURVEY OF THE MARINE FISHES OF FIJI

An intensive study of the marine fishes of Fiji was conducted over a five-year period (1999 – 2003). The many specimens collected during this survey are now in the CAS Ichthyology collection and are being identified. A number of species new to science have already been described.

A REVIEW OF THE GOBIES IN THE GENUS EVIOTA.

Species of Eviota are very small, with most species maturing at lengths less than 15 mm SL. Because of their small size and cryptic habits, these fishes are seldom seen by divers, but are one of the most abundant fishes in rotenone collections. These small gobies may form an important component of the food of larger piscivorous fishes. There are many species in this genus yet to be described.

TOADFISH SYSTEMATICS

I have an ongoing interest in toadfish systematics, specializing in Old World genera. My colleague, Dr. Bruce Collette of the Smithsonian Institution, and I have revised the genera and now are investigating several species-level problems.

Publications

1. Greenfield, D W. and R. B. Grinols. 1965. Cyprinodon variegatus on the Olympic Peninsula, Washington. Copeia 1965:115-116.

2. Miller, B. S. and D. W. Greenfield. 1965. A juvenile six-gilled shark (Hexanchus corinus) from the San Juan Islands, Washington. J. of the Fish. Res. Bd. of Canada 22:857-859.

3. Greenfield, D. W. 1965. Systematics and zoogeography of Myripristis in the eastern tropical Pacific Calif. Fish and Game 51:229-247.

4. Grinols, R. B. and D. W. Greenfield. 1966. New records of Cataetyx rubrirostris Gilbert from the northeastern Pacific Ocean. Calif. Fish and Game 52:211-213.

5. Greenfield, D. W. 1968. The zoogeography of Myripristis (Pisces. Holocentridae). Syst. Zool. 17:76-87.

6. Greenfield D. W. 1968. Observations on the behavior of the basket-weave cuskeel, Otophidium scrippsi Hubbs. Calif. Fish and Game 54:108-114.

7. Greenfield, D. W. and J. W. Wiley. 1968. Geographic variation in the clingfish, Gobiesox eugrammus Briggs. Trans. San Diego Soc. Nat. Hist. 15:141-147.

8. Greenfield D. W., D. A. Hensley, J. W. Wiley, and S. T. Ross. 1970. The fishes of the Isla Jaltemba coral formation and their zoogeographical significance. Copeia 1970:180-181.

9. Greenfield, D. W., S. T. Ross, and G. D. Deckert. 1970. Some aspects of the life history of the Santa Ana sucker, Catostomus (Pantosteus) santaanae (Snyder). Calif. Fish and Game 56:166-179.

10. Greenfield, D. W. and D. A. Hensley. 1970. Damselfishes (Pomacentridae) of Easter Island, with descriptions of two new species. Copeia 1970:689-695.

11. * Greenfield, D. W. 1971. Systematic Ichthyology: a collection of readings MSS Educational Publishing Co., New York, N.Y. 297 pp.

12. * Greenfield, D. W. 1971. The zoogeography of Myripristis (Pisces: Holocentridae) in relation to the Indo Australian Archipelago. Abstracts, Symposium on Indian Ocean and Adjacent seas, their origin, science and resources. Marine Biol. Assn. India, Cochin: 112-113. 590-592.

13. Greenfield, D.W. 1972. Notes on the biology of the arrow blenny, Lucayablennius zingaro (Böhlke) from British Honduras. Copeia 1972: 590-592.

14. Greenfield, D. W. and T. Greenfield. 1972. Introgressive hybridization between Gila orcutti and Hesperoleucus symmetricus (Pisces: Cyprinidae) in the Cuyama River Basin, California: I. Meristics, morphometrics and breeding. Copeia 1972:849-854.

15. Greenfield, D. W., F. Adbel-Hameed, G. D. Deckert and R. R. Flinn. 1973. Hybridization between Chrosomus erythrogaster and Notropis cornutus (Pisces: Cyprinidae). Copeia 1973:54-59.

16. Greenfield, D. W. and G. D. Deckert. 1973. Introgressive hybridization between Gila orcutti and Hesperoleucus symmetricus (Pisces: Cyprinidae) in the Cuyama River Basin, California: II. Ecological aspects. Copeia 1973:417-427.

17. Greenfield, D. W. and T. Greenfield. 1973. Triathalassothia gloverensis, a new species of toadfish from Belize (British Honduras), with remarks on the genus. Copeia 1973:560-565.

18. * Greenfield, D. W. 1973. Zoogeography of the coral-reef fishes of British Honduras. Amer. Philos. Soc. Yrbk. 1972:368-369.

19. Greenfield, D. W. 1973. An evaluation of the advisability of the release of the grass carp, Ctenopgharyngodon idella, into the natural waters of the United States Trans. Ill. State Acad. Sci. 66:47-53.

20. Greenfield, D. W. and L. P. Woods. 1974. Eupomacentrus diencaeus Jordan and Rutter, a valid species of damselfish from the western tropical Atlantic. Fieldiana Zool. 65:9-20.

21. Greenfield, D. W. 1974. A revision of the squirrelfish genus Myripristis Cuvier (Pisces: Holocentridae). Los Angeles Co. Mus., Sci. Bull. 19:1-54.

22. Thomerson, J. E. and D. W. Greenfield. 1975. Anchoviella belizensis, a new species of anchovy from Belize Central America, with records of associated freshwater species. Copeia 1975:50-52.

23. * Greenfield, D. W. 1975. (Review) Fishes of the World: A Key to Families and a Checklist. By G. U. Lindberg. Trans. Amer. Fish. Soc. 104:170-171.

24. Greenfield, D. W. 1975 Clepticus parrae, an additional sponge-dwelling fish. Copeia 1975: 381-382.

25. Greenfield, D. W. 1975. Centropomus poeyi from Belize with a key to the western Atlantic species of Centropomus. Copeia 1975:582-583.

26. Greenfield, D. W. 1975. Emblemariopsis pricei, a new species of chaenopsid from Belize. Copeia 1975:713-715.

27. Johnson, R. K. and D. W. Greenfield. 1976. A new chaenopsid fish, Emblemaria hyltoni, from Isla Roatan, Honduras. Fieldiana Zool. 70:13-28

28. Greenfield, D. W. and T. Greenfield. 1976. Fish collecting in Belize. Field Mus. Nat. Hist. Bull. 47:3-7, 15.

29. Greenfield, D. W. and T. Greenfield. 1977. The status of the Lake Maracaibo anchovy, Anchovia nigra Schultz (Pisces: Engraulidae). Brenesia 10/11:101-107.

30. * Greenfield, D. W. 1977. (Review) Investigations of the Ichthyofauna of Nicaraguan Lakes. By T. B. Thorson (ed.) Trans. Amer. Fish Soc. 106:196-197.

31. Greenfield, D. W. and G. S. Glodek. 1977. Trachelyichthys exilis, a new species of catfish (Pisces: Auchenipteridae) from Peru. Fieldiana Zool. 72:47-58.

32. * Greenfield, D. W. 1977. (Review) Fishes of the World. By J. S. Nelson. Trans. Amer. Fish. Soc. 106:408-409.

33. Woods, L. P. and D. W. Greenfield. 1978. Family Holocentridae. IN: FAO Species Idenification Sheets for Fishery Purposes, Western Central Atlantic, Vol. 3:1-10. W. Fisher (ed.).

34. Greenfield, D. W. 1979. A review of the Western Atlantic Starksia ocellata complex (Pisces: Clinidae) with the description of two new species and proposal of super-species status. Fieldiana Zool. 73:9-48.

35. Greenfield, D. W. and L. P. Woods. 1980. Review of the deepbodied species of Chromis (Pisces: Pomacentridae) from the eastern Pacific with descriptions of three new species. Copeia 1980:626-641.

36. Greenfield, D. W. and R. K. Johnson. 1981. The blennioid fishes from Belize and Honduras, Central America, with comments on their systematics, ecology, and distribution (Pisces: Labrisomidae, Chaenopsidae, Tripterygiidae, Blenniidae). Fieldiana Zool., New Ser. No.8:1-106

37. Greenfield, D. W. 1981. Varicus imswe, a new species of gobiid fish from Belize. Copeia 1981:269-272.

38. Greenfield, D. W. 1981. Family Holocentridae. IN: FAO Species Identification Sheets for Fishery Purposes, Eastern Central Atlantic, Vol. 2:1-10.

39. Johnson R. K. and D. W. Greenfield. 1982. Gone Fishing in the Gulf of Honduras. Field Mus. of Nat. Hist. Bull. 53:20-24.

40. Greenfield, D. W., T. A. Greenfield and D. M. Wildrick. 1982. The taxonomy and distribution of the species of Gambusia (Pisces: Poeciliidae) in Belize, Central America. Copeia 1982:128-147.

41. Greenfield, D. W. and T. A. Greenfield. 1982. Habitat and resource partitioning between two species of Acanthemblemaria (Pisces: Chaenopsidae), with comments on the chaos hypothesis: 499-507. IN: K. Rutzler and I.G. Macintyre (eds.) The Atlantic Barrier Reef Ecosystem at Carrie Bow Cay: Belize, 1: Structure and Communities. Smithsonian Contr. Mar. Sci. 12:539 pages. (1982).

42. Greenfield, D. W., T. A. Greenfield, and R. L. Woods. 1982. A new subspecies of cave-dwelling pimelodid catfish, Rhamdia laticauda typhla from Belize, Central America. Brenesia 19/20. 563-576.

43. Greenfield, D. W., C. F. Rakocinski, and T. A. Greenfield. 1983. Spatial and trophic interactions in wet and dry seasons between Gambusia luma and Gambusia sexradiata (Pisces: Poeciliidae) in Belize, Central America. Fieldiana, Zool. New Ser. No 14:1-16.

44. * Greenfield, D. W. and T. A. Greenfield. 1983. (Review) Peces Nicaraguenses de Agua Dulce. by J. Villa. Copeia 1983:282-283.

45. Johnson, R. K. and D. W. Greenfield. 1983. Clingfishes (Gobiesocidae) from Belize and Honduras, Central America, with a redescription of Gobiesox barbatulus Starks. Northeast Gulf Sci. 6:33-49.

46. Greenfield, D. W. 1983. Gambusia xanthosoma, a new species of poeciliid fish from Grand Cayman Island, B.W.I. Copeia 1983:457-464.

47. Greenfield, D. W., T. A. Greenfield, and S. L. Brinton. 1983. Spatial and trophic interactions between Gambusia sexradiata and Gambusia puncticulata yucatana (Pisces: Poeciliidae) in Belize, Central America. Copeia 1983:598-607.

48. Greenfield, D. W. and D. M. Wildrick. 1984. Taxonomic distinction of the Antilles Gambusia puncticulata complex (Pisces: Poeciliidae) from the G. yucatana complex of Mexico and Central America. Copeia 1984:921-933.

49. Greenfield, D. W. and J. D Rogner. 1984. An assessment of the fish fauna of Lake Calumet and its adjacent wetlands, Chicago, Illinois: past, present and future. Trans. Ill. State Acad. Sci. 77:77-93.

50. Greenfield, D. W. 1985. Review of the Gambusia yucatana complex (Pisces: Poeciliidae) of Mexico and Central America. Copeia 1985:368-378.

51. Wildrick, D. M., I. S. Kim, and D. W. Greenfield. 1985. A unique gambusine karyotype and its relevance to the Gambusia punctata species group. Copeia 1985:1053-1056.

52. Rakocinski, C. F. and D. W. Greenfield. 1985. Diet comparisons between savannah reed-pond Gambusia sexradiata and Gambusia puncticulata yucatana (deep-bodied form) (Pisces: Poeciliidae) on two dates during the dry season. Stud. Neotropical Fauna and Environ. 20(3):157-165.

53. Deckert, G. D and D W. Greenfield. 1987. A review of the western Atlantic species of the genera Diapterus and Eugerres (Pisces: Gerreidae). Copeia. 1987(1):182-194.

54. Greenfield, D. W. 1988. A review of the Lythrypnus mowbrayi Complex (Pisces: Gobiidae), with the description of a new species. Copeia. 1988(2):509-537.

55. Greenfield, D.W. 1989. Priolepis dawsoni n. sp. (Pisces: Gobiidae) a third Atlantic species. Copeia 1989 (2):397-401.

56. Greenfield, D.W. and R.K. Johnson. 1990. Community structure of western Atlantic Blennioid fishes. Copeia 1990(2):433-448.

57. Greenfield, D.W. 1990. Poecilia teresa, a new species of Poecilia from Belize, Central America Copeia 1990(2):449-454.

58. Greenfield, D.W. and R.K. Johnson. 1990. Heterogeneity in habitat choice in cardinalfish community structure. Copeia 1990(4):1077-1084.

59. * Greenfield, D.W. 1991. (Invited Review) Ecology and Evolution of live-bearing fishes (Poeciliidae). By Gary K. Meffe and Franklin F. Snelson. Copeia 1991(1):260-262.

60. Greenfield, D.W., L.T. Findley, and R.K. Johnson. 1993. Psilotris kaufmani n. sp. (Pisces: Gobiidae), a fourth western Atlantic species of Psilotris. Copeia 1993(1):183-186.

61. Greenfield, D. W. 1993. New goby, Psilotris boehlkei (Pisces: Gobiidae), from the western Atlantic, with a key to the species. Copeia 1993(3):771-775.

62. * Greenfield, D. W. 1994. Assemblage structure of small, cryptic, coral-reef fishes: an Atlantic-Pacific comparison. Proc. Fourth Indo-Pacific Fish Conference. Systematics and evolution of Indo-Pacific Fishes. Bangkok, Thailand: 338-339.

63. Greenfield, D. W., J. K. L. Mee, and J. E. Randall. 1994. Bifax lacinia, a new genus and species of toadfish (Batrachoididae) from the south coast of Oman. Fauna of Saudia Arabia 14: 276-281.

64. * Greenfield, D. W. 1995. Invited review - Proceedings of the colloquium on global aspects of coral reefs. American Zoologist 35(1):79-80.

65. Randall, J. E. and D. W. Greenfield. 1996. Revision of the Indo-Pacific holocentrid fishes of the genus Myripristis, with descriptions of three new species. Indo-Pacific Fishes, no. 25: 1-61.

66. Greenfield, D. W. 1996. Perulibatrachus kilburni, a new toadfish from East Africa (Teleostei: Batrachoididae). Copeia 1996(4): 901-904.

67. Greenfield, D. W. and J. E. Thomerson. 1997. The fishes of the continental waters of Belize. University Press of Florida, Gainesville. 330pp..

68. Greenfield, D. W. 1997. Allenbatrachus, a new genus of Indo-Pacific toadfish (Batrachoididae). Pacific Science 51(3): 306-313.

69. Greenfield, D. W., A. Y. Suzumoto and C. Chong. 1998. Vitraria clarescens, a junior synonym of the freshwater Hawaiian goby Sicyopterus stimpsoni (Teleostei: Gobiidae). Copeia 1998(2):501-503.

70. * Greenfield, D. W. 1998 Baseline studies of biodiversity: The fish resources of western Indonesia. D. Pauly and P. Martosubroto (eds). ICLARM. Quarterly Review of Biology 73(2):227.

71. Greenfield, D. W. 1998. Halophryne hutchinsi, a new toadfish (Batrachoididae) from the Philippine Islands and Pulau Waigeo, Indonesia. Copeia 1998(3): 696-701.

72. Greenfield, D. W. and R. K. Johnson. (1999). Assemblage structure and habitat associations of western Caribbean gobies (Teleostei: Gobiidae). Copeia 1999(2):251-266.

73. Greenfield, D. W. and J. E. Randall. (1999). Two new Eviota species from the Hawaiian Islands (Teleostei: Gobiidae). Copeia 1999(2):439-446.

74. Greenfield, D. W. (2000). Batrachoididae. In: FAO species identification guide for fishery purposes. The living marine resources of the Western Central Pacific, Vol. 3, Batoid fishes, chimaeras and bony fishes part 1:1999-2003. K. E. Carpenter and V. H. Niem (eds). FAO Rome.

75. Randall, J. E. and D. W. Greenfield (2000). Holocentridae. In: FAO species identification guide for fishery purposes. The living marine resources of the Western Central Pacific, Vol. 4, Bony fishes part 2:2225-2256. K. E. Carpenter and V. H. Niem (eds). FAO Rome.

76. * Greenfield, D. W. (2000). Obituaries. Robert Karl Johnson, 1944-2000. Copeia 2000 (4): 1148-1149.

77. Randall, J. E. and D. W. Greenfield. (2001). Preliminary review of the Indo-Pacific gobiid fishes of the genus Gnatholepis. Ichthyological Bulletin of the J. L. B. Smith Institute of Ichthyology, no. 69:1-17.

78. Greenfield, D. W. (2001). A revision of the Apogon erythrinus complex (Teleostei: Apogonidae). Copeia (2): 459-471.

79. Greenfield, D. W. (2001). Historical Retrospective: John E. Randall. Copeia 2001:872-877.

80. Greenfield, D. W. (2002). Palatogobius grandoculus, a new deep-water Western Caribbean goby (Teleostei: Gobiidae). Copeia 2002 (3):716-718.

81. Greenfield, D. W., and K. Matsuura. (2002). Scorpaenodes quadrispinosus, a new Indo-Pacific scorpionfish (Teleostei: Scorpaenidae). Copeia 2002 (4):973-978.

82. Greenfield, D. W. (2003). A survey of the small reef fishes of Kaneohe Bay, Oahu, Hawaiian Islands. Pacific Science 57(1):45-76.

83. Greenfield, D. W. (2003). Review. Coral reef fishes: Dynamics and diversity in a complex ecosystem. Copeia 2003 (1):214-216.

84. Greenfield, D. W. (2003). Pacific-Atlantic differences in cardinalfish assemblage structure. Copeia 2003 (3):608-617.

85. Greenfield, D. W. (2003). Holocentridae. In: The living marine resources of the Western Central Atlantic. FAO Species Identification guide for Fishery Purposes and Amer. Soc. Ichthyologists and Herpetologists Special Publ. No. 5, vol. 2, pt. 1:1192-1202.

86. Gilmore, R. G., Jr. and D. W. Greenfield. (2003). Gerreidae. The living marine resources of the Western Central Atlantic. FAO Species Identification Guide for Fishery Purposes and Amer. Soc. Ichthyologists and Herpetologists Special Publ. No. 5, vol 3, pt. 2:1506-1516.

87. Greenfield, D. W. and S. Neisz. (2004). Gorgasia thamani, a new species of garden eel from Fiji (Teleostei: Congridae; Heterocongrinae). Proceedings California Academy of Sciences, 55 (17): 371-374.

88. Randall, J. E. and D. W. Greenfield (2004). Two new scorpionfishes (Scorpaenidae) from the South Pacific. Proceedings California Academy of Sciences, 55 (19):382-392.

89. Greenfield, D. W. and J. E. Randall. 2004. The marine gobies of the Hawaiian Islands. Proceedings California Academy of Sciences, 55(27):500-551.

90. Greenfield, D. W. and Wm. L. Smith. 2004. Allenbatrachus meridionalis, a new toadfish (Batrachoididae) from Madagascar and Reunion. Proceedings California Academy of Sciences, 55(28):568-572.

91. Greenfield, D. W. and J. E. Randall. 2004. Two new cardinalfish species of the genus Apogon from Easter Island. Proceedings California Academy of Sciences, 55(29):563-569.

92. Wilk, K., K. Wilk, and D. W. Greenfield. 2005. First record of the Eastern Atlantic squirrelfish Sargocentron hastatum (Holocentridae) from the Western Atlantic Ocean. Proceedings California Academy of Sciences, 56-SC:86-87.

93. Greenfield, D. W. 2005. Perulibatrachus aquilonarius, a new toadfish species from India (Teleostei: Batrachoididae). Proceedings California Academy of Sciences, 56 (7):76-79.

94. Greenfield, D. W. 2005. Brotula flaviviridis, a new species of brotula from Fiji (Teleostei: Ophidiidae: Brotulinae). Proceedings Callifornia Academy of Sciences, 56 (8):80-85.

95. Greenfield, D.W. and K. R. Longenecker. 2005. Vanderhorstia bella, a new goby from Fiji (Teleostei: Gobiidae). Proceedings California Academy of Sciences, 56 (32):619-623.

96. Greenfield, D.W., R. C. Langston, and J. E. Randall. 2005. Two new cardinalfishes of the Indo-Pacific fish genus Zoramia (Apogonidae). Proceedings California Academy of Sciences, 56 (33):625-637.

97. Greenfield, D.W. 2006. Reef and shore fishes of the South Pcific: New Caledonia to Tahiti and the Pitcairn Islands (Review). Copeia 2006 (2):322-323.

98. Collette, B.B., D.W. Greenfield, and J. L. Costa 2006. Are there one or two species of Halobatrachus toadfishes (Teleostei: Batrachoididae) in tahe eastern Atlantic? Proceedings California Academy of Sciences, 57 (31): 937-943.

99. Greenfield, D.W. 2006. Two new toadfish genera (Teleostei: Batrachoididae). Proceedings California Academy of Sciences, 57 (32): 945-954

100. Randall, J.E. and D.W. Greenfield 2007. Redescription of Gobius cauerensis Bleeker, with discussion of the validity of the species. Zool. Med. Leiden 81:303-308.

101. Greenfield, D.W. 2007. Apogon seminigracaudus, a new cardinalfish species previously misidentified as Apogon fuscus (Teleostei: Apogonidae). Proceedings California Academy of Sciences, Fourth Series, 58(17):361-366.

102. Greenfield, D.W. 2007. Reef and shore fishes of the Hawaiian Islands (Review). Copeia 2007(3): 775-777.

103. Greenfield, D.W. 2007. Geographic variation in a cardinalfish, Apogon dianthus (Teleostei: Apogonidae). Proceedings of the California Academy of Sciences, Fourth Series, 58(30): 601-605.

104. Greenfield, D.W. and J.E. Randall 2008. Eviota tigrina, a new goby from Tonga (Teleostei: Gobiidae). Proceedings of the California Academy of Sciences, Fourth Series, 59(11): 497-501

105. Greenfield, D.W., R. Winterbottom, and B.B. Collette. 2008. Review of the toadfish genera (Teleostei: Batrachoididae). Proceedings of the California Academy of Sciences, Fourth Series, 59(15): 665-710

106. Collette, B.B. and D.W. Greenfield. 2009. Batrachus uranoscopus Guichenot, 1866 supposedly from Madagascar, is not a threatened species of toadfish (Batrachoididae). Cybium 33(1):79-80.

107. Greenfield, D.W. 2009. Eviota randalli, a new gobiid fish from Oceania. Proceedings of the California Academy of Sciences, Fourth Series, 60(20):683-687.

108. Greenfield, D.W. and J.E. Randall 2010. Four new gobiid fishes of the genus Eviota from the Western Pacific, with clarification of Eviota guttata and Eviota albolineata (Teleostei: Gobiidae). Proceedings of the California Academy of Sciences, Fourth Series, 61(3):269-289.

109. Greenfield, D.W. and T. Suzuki. 2010. Eviota nigrispina, a new goby from the Ryukyu Islands, Japan (Teleostei: Gobiidae). Zootaxa 2655: 57-62.

110. Greenfield, D.W. and J.E. Randall 2010. Eviota karaspila, a new gobiid fish from Fiji (Teleostei: Gobiidae). Zootaxa 2672: 61-68.

111. Greenfield, D.W. and T. Suzuki. 2011. Two new goby species of the genus Eviota from the Ryukyu Islands, Japan (Teleostei: Gobiidae). Zootaxa 2812:63-68.

112. Bineesh, K.K. and D.W. Greenfield. 2011. A redescription of the Indian toadfish Perulibatrachus aquilonarius Greenfield (Teleostei: Batrachoididae). Zootaxa:2886:63-68.

113. Greenfield, D.W. and J.E. Randall. 2011. Two new Indo-Pacific species in the Eviota nigriventris complex (Teleostei: Gobiidae). Zootaxa: 2997:54–66.

114. Greenfield, D.W. and S.L. Jewett. 2011. Eviota rubriceps, a new goby from the southwestern Pacific Ocean, with comments on E. mikiae and E. raja (Teleostei: Gobiidae). Zootaxa 3134:53-62.

115. Greenfield, D.W. 2012. Colletteichthys occidentalis, a new toadfish species from the Arabian peninsula and northern Arabian Sea (Teleostei: Batrachoididae). Zootaxa 3165:64–68.

116. Greenfield, D.W. and T. Suzuki. 2012. Eviota atriventris, a new goby previously misidentified as Eviota pellucida Larson (Teleostei: Gobiidae). Zootaxa 3197: 55-62.

117. Greenfield, D.W. 2012. Shore fishes of Easter Island. (Review). Copeia.2012(1):165-166.

118. Greenfield, D.W. 2012. Austrobatrachus iseleseli, a new toadfish species from South Africa (Teleostei: Batrachoididae). Zootaxa 3400: 58–63.

119. Greenfield, D.W., K.K. Bineesh, and K.V. Akhilesh. 2012. Colletteichthys flavipinnis, a new toadfish species from Sri Lanka and India (Teleostei: Batrachoididae). Zootaxa 3437: 24–31.

120. Greenfield, D.W. and S.L. Jewett. 2012. Two new gobiid fishes of the genus Eviota from the Indian Ocean (Teleostei: Gobiidae). Zootaxa 3515: 67–74.

121. Greenfield, D.W. and G.R. Allen 2012. Eviota fallax, a new dwarf goby from the Western Pacific (Teleostei: Gobiidae). Zootaxa.3522: 42-48.

122. Greenfield, D.W. and R. Winterbottom. 2012. Two new dwarf gobies from the Southwestern Pacific Ocean (Teleostei: Gobiidae: Eviota). Zootaxa 3572: 33-42

123. Greenfield, D.W. and T. Suzuki. 2013. Eviota nigramembrana, a new dwarfgoby from the Western pacific (Teleostei: Gobiidae). Zootaxa 3637 (2): 169–175.

124. Greenfield, D.W. and M.V. Erdmann. 2013. Eviota santanai, a new Dwarfgoby from Timor-Leste (Teleostei: Gobiidae). Zootaxa 3741 (4): 593–600.

125. Greenfield, D.W, Bogorodsky, S.V., and Mal, A.O. 2014. Two new Red Sea dwarfgobies (Teleostei, Gobiidae, Eviota). Journal of the Ocean Science Foundation 10: 1–10.

126. Greenfield, D.W. and R. Winterbottom. 2014 Eviota piperata, a new gobiid species from Palau (Teleostei: Gobiidae). Zootaxa, 3755 (3): 295-300.

127. Greenfield, D.W., T. Suzuki, and K. Shibukawa. 2014. Two new dwarfgobies of the genus Eviota from the Ryukyu Islands, Japan (Teleostei: Gobiidae). Zootaxa, 3774 (5): 481–488.

128. Greenfield, D.W., R. Winterbottom, and T. Suzuki. 2014. Eviota occasa a new species of dwarfgoby from Palau and the Ryukyu Islands, Japan (Teleostei: Gobiidae). Journal of the Ocean Science Foundation, 10: 11–19.

129. Greenfield, D.W. and L. Tornabene. 2014..Eviota brahmi n. sp. from Papua New Guinea, with a redescription of Eviota nigriventris (Teleostei: Gobiidae). Zootaxa, 3793 (1):133-146.

130. Greenfield, D.W. and S.L. Jewett. 2014. Two new dwarfgobies from the Western Pacific (Teleostei: Gobiidae: Eviota). Copeia, 2014 (1): 56–62.

131. Greenfield, D.W. and M.V. Erdmann. 2014. Eviota tetha, a new species of dwarfgoby from Cendrawasih Bay, Papua, Indonesia (Teleostei: Gobiidae). Journal of the Ocean Science Foundation, 11 23-31.

132. Suzuki, T. and D.W. Greenfield. 2014. Two new dwarfgobies from the Ryukyu Islands, Japan: Eviota shibukawai and Eviota filamentosa (Teleostei: Gobiidae). Journal of the Ocean Science Foundation, 11: 32-39.

133. Greenfield, D.W. and S.L. Jewett. 2014. Eviota minuta, a new dwarfgoby from the Philippine Islands (Teleostei: Gobiidae). Journal of the Ocean Science Foundation 12: 12-17.

134. Greenfield, D.W. and M.V. Erdmann. 2014. Eviota algida, a new dwarfgoby species from the upwelling waters off Nusa Penida, Indonesia (Teleostei: Gobiidae). Journal of the Ocean Science Foundation 13: 27-34.

135. Greenfield, D.W. 2014. Barchatus indicus, a new toadfish species from Somalia (Teleostei: Batrachoididae). Copeia 2014(4): 668-672..

136. Suzuki, T., Greenfield, D.W., and Motomura, H. 2015. Two new dwarfgobies (Teleostei: Gobiidae) from the Ryukyu Islands, Japan: Eviota flavipinnata and Eviota rubrimaculata. Zootaxa 4007 (3): 399-408.

137. Greenfield, D.W., and Randall, J.E. 2016. A review of the dwarfgobies of Fiji, including descriptions of five new species (Teleostei: Gobiidae: Eviota). Journal of the Ocean Science Foundation 20: 25–75.

138. Greenfield, D.W., and Jewett, S.L. 2016. Two new dwarfgobies from the Indian and Western Pacific Oceans (Teleostei: Gobiidae: Eviota). Zootaxa 4121 (5): 580–599.

139. Greenfield, D.W., and Jewett, S.L. 2016. Eviota melanosphena, a new dwarfgoby from Australia (Teleostei: Gobiidae). Journal of the Ocean Science Foundation 2016 (21): 71–77.

140. Tornabene, L. & Greenfield, D.W. (2016) Eviota erdmanni (Teleostei: Gobiidae), a new dwarfgoby from the Savu Sea, Flores, Indonesia. Journal of the Ocean Science Foundation 2016 (22): 1–9.

141. Greenfield, D.W, & Winterbottom, R. (2016) Three new dwarfgobies from the western Pacific (Teleostei: Gobiidae: Eviota). Journal of the Ocean Science Foundation 2016 (22): 28-40.

142. Collette, B. B, and D. W. Greenfield (2016). Batrachoididae. In: Carpenter, K., P. Oliver, M. Lamboeuf, and J. Martin (eds.). The living marine resources of the Eastern Central Atlantic. FAO Species Identification Guide for Fishery Purposes. Rome, FAO, vol. 3: 2036-2043.

143. Greenfield, D. W. (2016). Holocentridae. In: Carpenter, K., P. Oliver, M. Lamboeuf, and J. Martin (eds.). The living marine resources of the Eastern Central Atlantic. FAO Species Identification Guide for Fishery Purposes. Rome, FAO, vol. e: 2196-2202..

144. Greenfield, D.W.and R. Winterbottom (2016). Eviota sodwanaensis, a new dwarfgoby from South Africa (Teleostei: Gobiidae). Journal of the Ocean Science Foundation 2016 (22): 53-57.

145. Greenfield, D.W., Suzuki, T. (2016). Eviota bilunula, a new dwarfgoby species from Fiji, with a redescription of Eviota flebilis (Teleostei: Gobiidae). Journal of the Ocean Science Foundation 2016 (24): 1-9.

146. Greenfield, D.W. (2016). Batrachoididae. In: Psomadakis, P.N., Osmany, H.B., & Moazzam, M. (eds.).Field identification guide to the living marine resources of Pakistan. FAO Species Identification Guide for Fishery Purposes. Rome. 174.

147. Greenfield, D.W. & Winterbottom, R. (2016). A key to the dwarfgoby species (Teleostei: Gobiidae: Eviota) described between 1871 and 2016. Journal of the Ocean Science Foundation 2016 (24): 35-90.

148. Greenfield, D.W. & Randall, J.E. (2017). Sueviota pyrios, a new species of coral-reef goby from the Red Sea (Teleostei: Gobiidae). Journal of the Ocean Science Foundation 2017 (25): 8-13.

149. Greenfield, D.W., Randall, J.E., & Psomadakis, P.N. (2017). A review of the soldierfish genus Ostichthys (Beryciformes: Holocentridae), with descriptions of two new species from Myanmar. Journal of the Ocean Science Foundation 2017 (26): 1-33.

150. Greenfield, D.W, Tornabene, L., & Erdmann, M.V. (2017). Eviota imitata, a new dwarfgoby from Raja Ampat, Indonesia (Teleostei: Gobiidae). Journal of the Ocean Science Foundation 2017 (26): 86-94.

151. Greenfield, D.W. & Erdmann, M.V. (2017). Eviota pictifacies, a new dwarfgoby from Sumbawa, Indonesia (Teleostei: Gobiidae). Journal of the Ocean Science Foundation 2017 (27): 1-7.

152. Buchanan, J.R.,Carpenter, K.E.,Greenfield, D.W.,Harvey, M.S.,Harwell, H.D.,Larson, H.K.,Lawrence, A.,Linardich, C.,Matsuura, K.,Motomura, H.,Munroe, T.A.,Myers, R.F.,Pippard, H.,Ralph, G.M.,Russell, B.C.,Smith-Vaniz, W.F.,Thaman, R.R.,Vié, J.-C.,Williams, J.T. (2017). The conservation status of marine biodiversity in the Pacific Islands of Oceania. International Union for Conservation of Nature and Natural Resources, 59pp.

153. Greenfield, D.W. (2017). Tomiyamichthys reticulatus, a new species of shrimpgoby from Fiji (Teleostei: Gobiidae). Journal of the Ocean Science Foundation 2017 (28): 47-51.

154. Greenfield, D.W. (2017) An overview of the dwarfgobies, the second most speciose coral-reef fish genus (Teleostei: Gobiidae: Eviota). Journal of the Ocean Science Foundation 2017 (29): 32-54.

155. Greenfield, D.W. & Randall, J. E. 2017. Eviota lentiginosa, a new dwarfgoby from Norfolk Island, Australia (Teleostei: Gobiidae). Journal of the Ocean Science Foundation 2017 (29): 66-71.

156. Greenfield, D.W. & Allen, G.R. (2018) Cryptocentrus nanus, a new species of dwarf shrimpgoby from Fiji (Teleostei: Gobiidae). Journal of the Ocean Science Foundation 2018 (30): 28-38.

157. Greenfield, D.W. & Randall, J.E. (2018) Myersina balteata, a new shrimp-associated goby (Teleostei: Gobiidae) from Guadalcanal, Solomon Islands. Journal of the Ocean Science Foundation 2018 (30): 90-99.

158. Greenfield, D.W., Tornabene, L., Gomez-Buckley, M., & Erdmann, M.V. (2018) Eviota maculosa, a new dwarfgoby from the Western Pacific Ocean (Teleostei: Gobiidae). Journal of the Ocean Science Foundation 2018 (31): 18-31.

159. Greenfield, D.W., Tornabene, L., Erdmann, M.V., & Pada, D.N. (2019) Eviota gunawanae, a new microendemic dwarfgoby from the Fakfak Peninsula, West Papua, Indonesia (Teleostei: Gobiidae). Journal of the Ocean Science Foundation 2019 (32): 57-67.

160. Greenfield, D.W. & Gordon, L. (2019) Eviota dalyi, a new dwarfgoby from the Amirante islands, Seychelles (Teleostei: Gobiidae). Journal of the Ocean Science Foundation 2019 (33): 9-15.

161. Greenfield, D.W. & Erdmann, M.V. (2019) Redescription of Eviota thamani, an endemic dwarfgoby from Fiji (Teleostei: Gobiidae). Journal of the Ocean Science Foundation 2019 (33): 65-69.

162. Greenfield, D.W., Erdmann, M.V., & Utama, I.V. (2019) Sueviota minersorum, a new species of sponge-dwelling goby (Teleostei: Gobiidae) from Misool, Raja Ampat Islands, Indonesia. Journal of the Ocean Science Foundation 2019 (33): 79-88.

163. Winterbottom, R. & Greenfield, D.W. (2020). Eviota pseudaprica, a new dwarfgoby from the Western Pacific Ocean (Teleostei: Gobiidae). Journal of the Ocean Science Foundation 2020 (35): 30-40.

164. Greenfield, D.W. & Erdmann, M.V. (2020). Eviota marteynae, a new dwarfgoby from the Maldive islands (Teleostei: Gobiidae). Journal of the Ocean Science Foundation 2020 (35): 94-101.

165. Greenfield, D.W. & Erdmann, M.V. (2020). Eviota amphipora, a new dwarfgoby fromn Papua New guinea (Teleostei: Gobiidae). Journal of the Ocean Science Foundation 2020 (36): 1-5.

166. Greenfield, D.W. & Erdmann, M.V. (2020). Eviota angustifascia, a new dwarfgoby from Fiji and New Guinea (Teleostei: Gobiidae). Journal of the Ocean Science Foundation 2020 (36): 31-37.

167. Greenfield, D.W. (2021). Addendum to the 2016 key to the dwarfgoby species (Teleostei: Gobiidae: Eviota). Journal of the Ocean Science Foundation 2021 (38): 1-12.

168. Greenfield, D.W. & Erdmann, M.V. (2021). Eviota flaviarma, a new dwarfgoby from Papua New Guinea (Teleostei: Gobiidae). Journal of the Ocean Science Foundation 2021 (38): 1-27-34.

169. Tornabene, L, Greenfield, D.W. & Erdmann, M.V. (2021). A review of the Eviota zebrina complex, with the description of four new species (Teleostei: Gobiidae). Zookeys 1057:149-184.

170. Greenfield, D.W. (2022). Batrachoididae. Coastal fishes of the Western Indian Ocean. P.C. Heemstra, P.C. et al.(eds.) Vol. 2, 271-282, plates 44-46. South Africa Institute for Aquatic Diversity, Makhanda, South Africa.

171. Greenfield, D.W., Erdmann, M.V., & Mambrasar, R. (2022). Eviota fluctiphila, a new dwarfgoby from the western Pacific Ocean (Teleostei: Gobiidae). Journal of the Ocean Science Foundation 2022 (39): 36-44..

172. Erdmann, M.V., Greenfield, D.W., & Tornabene, L. (2023). Eviota taeiae, a new dwarfgoby (Teleostei: Gobiidae) of the Eviota guttata complex from Samoa. Journal of the Ocean Science Foundation 2023 (40): 37-47.

173. Greenfield, D.W., Erdmann, M.V., & Tornabene, L. (2023). Eviota atauroensis, a new dwarfgoby (Teleostei: Gobiidae) from Timor-Leste. Journal of the Ocean Science Foundation 2023 (40): 48-57.

 

* Not peer reviewed.

The World Book Encyclopedia:

1988 Flounder; Minnow; Muskellunge; Shad; Sturgeon; Trout; Turbot

1990 Bullhead; Roach; Smelt; Stickleback; Whitefish

1991 Eulachon; Grayling; Sucker

1997 Salmon

1998 Capelin

2003 Haddock, Halibut, Herring

2005 Goldfish

Grzimek’s Animal Life Encyclopedia - 2003, 2nd Edition, Vol. 5, Fishes II: 41-45.

Batrachoidiformes: Toadfishes

Maritimea: Above and Beneath the Waves – 2009, Millennium House, Australia: Fishes, pg. 182-201.

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CV

Education

Ph.D. 1966 (Fisheries) University of Washington

B.A. 1962 (Zoology) California State University, Humboldt

Areas of Specialization

 

Ichthyology, Marine Biology, Biosystematics, Zoogeography, Community Ecology

 

Administrative Experience

1995. President, American Society of Ichthyologists and Herpetologists

1993 - 1994. President, Western Association of Graduate Schools

1991 - 1995. Executive Committee, Western Association of Graduate Schools

1987 - 1995. Dean of The Graduate Division, University of Hawaii at Manoa

1986 - 1987. Associate Vice Chancellor for Academic Affairs and Dean of the Graduate School, University of Colorado at Denver

1986. Acting Vice Chancellor for Academic Affairs, University of Colorado at Denver (7 1/2 months)

1984 - 1986. Associate Vice Chancellor for Academic Affairs and Dean of the Graduate School, University of Colorado at Denver

1984. Interim Dean of the Graduate School and Associate Provost for Graduate Studies and Research, Northern Illinois University

1982 - 1984. Associate Dean, Graduate School, Northern Illinois University

1979 - 1981. Program Director: Coral-Reef Biology, Belize, Central America

1972 - 1973. Director: A.U.I.E. Tropical Studies Center, Belize, Central America

Teaching Experience

2003 - present. Emeritus Professor of Zoology, University of Hawaii at Manoa

1987 - 2003. Professor of Zoology, University of Hawaii at Manoa

1984 - 1987. Professor of Biology, University of Colorado-Denver

1977 - 1984. Professor of Biological Sciences, Northern Illinois University, DeKalb, Illinois

1977 (summer). Visiting Faculty, Friday Harbor Laboratories, University of Washington, Friday Harbor, Washington

1975 - 1976. Sabbatical leave, St. John's College, Belize, Central America and Hopkins Marine Station, Stanford University

1971 - 1975. Marine Faculty, A.U.I.E. Tropical Studies Center, Belize Central America

1970 - 1977. Associate Professor, Northern Illinois University, DeKalb, Illinois

1966 - 1970. Assistant Professor, California State University, Fullerton, California

1965 (summer). Instructor, NSF Radiation Biology Summer Institute, University of Washington, Seattle, Washington

1962 - 1964. Teaching Assistant, University of Washington, Seattle, Washington

Other Professional Experience

2004 - present. Adjunct Research Associate, Natural History Museum and Biodiversity Research Center, University of Kansas

2003 - present. Research Affiliate, Moss Landing Marine Laboratory

2002 - present. Research Associate in Ichthyology, California Academy of Sciences

1991 - present. Research Associate in Zoology, Bishop Museum, Honolulu, Hawaii

1991 - 2003. Affiliate Researcher, Hawaii Institute of Marine Biology

1974 - 1991. Member Board of Directors, ENCAP, Inc., Environmental Consultants and Planners, DeKalb, Illinois

1973 - present. Research Associate Division of Fishes. Field Museum of Natural History, Chicago, Illinois

1964 - 1966. Research Assistant, University of Washington

1963 (summer) Research Assistant, U S. Fish and Wildlife Service, San Diego, California Professional Activities:

1995. President, American Society of Ichthyologists and Herpetologists

1987 - 1992. Member, Board of Governors, American Society of Ichthyologists and Herpetologists

1983 - 1984. Member, Nominating Committee, American Society of Ichthyologists and Herpetologists

1982. Local Chair, 62nd Annual Meeting of the American Society of Ichthyologists and Herpetologists, DeKalb, Illinois, June 1982

1980 - 1985. Member, Board of Governors, American Society of Ichthyologists and Herpetologists

1979 - 1983. Member, Time and Place Committee, American Society of Ichthyologists and Herpetologists

1978. Participant, Virgin Islands Offshore Sand Dredging Workshop, St. Thomas, Virgin Islands, NOAA

1974 - 1975. Member of Committee on Environmental Quality, American Society of Ichthyologists and Herpetologists

1973 - 1978. Member, Board of Governors, American Society of Ichthyologists and Herpetologists

1971. Participant, International Decade of Oceanic Exploration, C.I.T.R.E. Workshop, Belize

1969 - 1970. Member, American Fisheries Society and American Society of Ichthyologists and Herpetologists Joint Committee on Introduction of Exotic Fishes and other Aquatic Organisms

1969 - 1970. Index Editor for Copeia, American Society of Ichthyologists and Herpetologists

Professional Organizations

American Society of Ichthyologists and Herpetologists

Honors:
2005. Fellow of the California Academy of Sciences

1975. Fulbright-Hays Grant for Colombia, South America- Awarded but declined due to conflicting N.S.F. Grant

1965 - 1966. Research Fellowship, Graduate School, University of Washington, Seattle, Washington

1964. Stoye Prize, American Society of Ichthyologists and Herpetologists-best student paper.

Research Support

2001. National Science Foundation. A survey of the marine fishes of Fiji.

2001. National Geographic Society. The impact of coral bleaching on the coral- reef fish community.

2001. Sea Grant. Characterizing the role of small, cryptic fishes in the trophic structure of coral-reef ecosystems.

1999. Sea Grant. Assemblage structure of small, cryptic fishes in coral reefs and adjacent habitats—a pilot baseline study for protected area designation in Fiji.

1998. National Science Foundation: Planning visit for a study of the cryptic fish communities at Fiji

1995. Sea Grant: Assemblage structure and trophic relationships of the spur and groove cryptic fish community

1985. National Science Foundation: A survey of the Cuban shorefish fauna.

1980. National Science Foundation: Systematics of the Gambusia puncticulata- yucatana species complex

1980. Graduate School, Northern Illinois University: Distribution and habitat associations of coral-reef fishes of the northern Barrier Reef in Belize, Central America.

1975. National Science Foundation: Isolating mechanisms and competitive interactions of gambusiine fishes of Belize.

1974. Pet Industry Joint Advisory Council: Establishment of Aquarium Sciences program.

1973. U S. Bureau of Sport Fisheries and Wildlife: Impact of commercial collecting on wild-caught ornamental aquarium fishes of South America. Contract participant.

1973. The Council of Academic Deans, Northern Illinois University, DeKalb, Illinois: An investigation of the fish fauna of the mangrove and sea grass habitats of British Honduras.

1970. The Council of Academic Deans, Northern Illinois University, DeKalb, Illinois: Zoogeography of the coral-reef fishes of British Honduras.

1970. American Philosophical Society: Zoogeography of the coral-reef fishes of British Honduras.

1970. Faculty Research Award, California State University, Fullerton: The ecology of Chromis.

1967. California Partners of the Alliance (Mexico): A survey of the fishes of the coast of Nayarit.

1965. Sigma Xi Grant-In-Aid of Research: Systematics of Myripristis.

1965. Karl P. Schmidt Research Grant, Field Museum of Natural History, Chicago, Illinois: Systematics of Myripristis.

Institutional Grants

1988,1990, 1991,1992. U.S. Department of Education: Encouragement of minority participation in graduate education

1991. Patricia Roberts Harris Fellowships Program.

 

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About the Ichthyology Department

Learn more about the Ichthyology department.

Ichthyology Research Associates

Daniel M. Cohen
Research Associate

David A. Ebert
Program Director / Adjunct Faculty
Pacific Shark Research Center, Moss Landing Marine Laboratories

Mark Van N. Erdmann
Senior Advisor
Conservation International Indonesia Marine Program

Carl J. Ferraris
Research Associate

Michelle R. Gaither
Research Associate

David W. Greenfield
Research Associate
Emeritus Professor, Univ. of Hawaii

Robert N. Lea
Research Associate

Douglas J. Long
Research Associate

Catherine W. Mecklenburg
Point Stephens Research

Rita S. Mehta
Assistant Professor University of California, Santa Cruz

David A. Neely
Research Associate

William J. Poly
Research Associate

Stuart G. Poss
Research Associate

Graham Short
Research Associate 

Ichthyology at the Academy

The Department of Ichthyology is home to one of the largest and most important collections of fishes in the world, and is designated as one of eight International Centers for Ichthyology in North America. Meet the researchers, explore projects and expeditions, and more.